Depending on how the gravel's being used, that can happen
![Smile :) :)](data:image/gif;base64,R0lGODlhAQABAIAAAAAAAP///yH5BAEAAAAALAAAAAABAAEAAAIBRAA7)
. However, if the inlet for the filter is underneath the gravel bags, then the amount of surface area for bacterial growth for the cycle is increased, and particulate waste can be caught, then if appropriate organisms such as blackworms are included in the system, utilised by them or by plants.
Also, I would disagree. Just because it hasn't happened doesn't mean it won't (more it's a matter of when it will happen).
Worse, someone will see the zoo's axolotls and think "hey, gravel works!", set up their new tank at home and be feeding them sinking pellets or something else that can work into the rocks, and we end up with them on here in tears because their beloved new pet is dying.
The role of a zoo is to provide extremely high standards of care and husbandry for their animals so that visitors are able to see these animals in the best condition they can be, and to ensure that they conserve animals in a manner that is appropriate for the species they are dealing with. As a community, we have seen far too much evidence of the dangers of gravel, so to hear of a "professional" conservation organisation ignoring the insurmountable evidence is extremely disconcerting and needs to be dealt with both for the safety of the axolotls and for the ongoing standards of care provided to axies around the world.
And please don't take this as me going off at you- I'm angry at that zoo for not providing an appropriate habitat despite obviously knowing what the evidence says, and harbouring a much deeper anger and frustration with the pet shops and dealers who refuse to observe proper husbandry practices for their animals, especially axies who by far and away seem to take the brunt of malpractice.